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Borland, 24, said he notified the 49ers on Friday. He said he made his decision after consulting with family members, concussion researchers, friends and current and former teammates, as well as studying what is known about the relationship between football and neurodegenerative disease.

“We’ve been on the front lines,” Dubois says, “of accepting and being willing to talk about our exploration.”

Tramaine Brock was at left cornerback on the first string and backed up by Dontae Johnson, as well as rookie Rashard Robinson. Kenneth Acker and Keith Reaser saw action on the right side.

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